The game begins with Reed, Sue, and Johnny on a roof lying helpless after being knocked unconscious by a blast from Doctor Doom. Sue is the first to recover, so when she sits up and turns around, Dr. Doom is preparing to fire an electric blast at them. Sue holds it off with her force field and calls Ben Grimm for help. Then it cuts away to Ben, who is recovering in the transformation chamber after having his rocklike exterior genetically removed from him. As he contemplates why the circumstances have led to this, he remembers the period when all of this happens.
This is when it reverts to the beginning of the movie: Reed signs the pact, they go into space, and Ben gets the samples ready. In space, they are hit by a cosmic storm which alters their DNA and gives them superpowers. If it hadn't been for Victor, they might never have arrived back to Earth and into his medical compound, where they recovered. When Ben discovers that he has become a monster-like figure, he deserts the other three and heads home.
Ben runs on a rampage to try to calm himself.
This brings the army to New York under the control of a deep, dark, sinister figure who is yet to be revealed and they try to get Ben under control. However, after Ben and the other three rescue a fire truck from falling off the Brooklyn Bridge, the forces lay off and watch the Fantastic Four to see if they become hostile.
Reed attempts to find another power source, but is interrupted by a call for help. It seems that strange creatures have invaded Grand Central Station and the police seem to have no effect against them. The Fantastic Four stop the creatures from invading the city and face their leader, the Mole Man and his mighty pet. Because of the utter destruction caused by their fight with this giant monster, the city is in a mess and Victor blames Reed for all this mess
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